How exactly does 401k work?

How exactly does 401k work? I've just been curious. So I know companies do matching up to 6% or so. But those matching funds require vesting, idk maybe 2 or3 years. If you leave they take them back. But isnt the 401k a brokerage account? The money is used to buy some combination of stocks and bonds. Due to this the value of the account can fluctuate. So I don't get it. So ...

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